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Carrying Capacity

Created Dec 23, 2024 ecologylimitssystems

Carrying capacity (often denoted K) is the maximum population size that an environment can sustain indefinitely given available resources. Above this level, mortality exceeds reproduction and population declines. Below it, population grows. At carrying capacity, the system is in equilibrium.

The concept comes from the logistic growth model, developed by Pierre François Verhulst in 1838. Population growth slows as it approaches carrying capacity, producing an S-curve: exponential at first, then decelerating, then leveling off. The model fits some populations well and others poorly, but the underlying concept (that environments have limits) is fundamental.


Carrying capacity isn’t fixed. It depends on technology, behavior, and environmental conditions. Agricultural intensification raised human carrying capacity. Drought lowers carrying capacity for herbivores. A species that learns to exploit a new food source effectively increases K for itself. The limit is real but movable.

Different resources create different limits. A population might have ample food but limited nesting sites. Water might limit desert populations, nitrogen might limit temperate forests. The scarcest resource relative to need sets the effective carrying capacity — Liebig’s law of the minimum.


Populations can exceed carrying capacity temporarily, but not sustainably. Overshoots happen when reproduction continues faster than feedback signals the limit. The population then crashes, sometimes below original levels because overshoot degrades the resource base. Deer overpopulating an island strip vegetation; the crash comes when winter arrives with no forage.

The pattern appears in many domains. Market bubbles exceed sustainable valuations. Cities grow faster than infrastructure. Debts accumulate faster than repayment capacity. Overshoot followed by correction is a general systems pattern, not specific to ecology.


Human carrying capacity remains contested. Malthus predicted population would outstrip food in the late 18th century; it hasn’t yet. The Green Revolution expanded agricultural output. Demographers expect global population to peak at 10-11 billion this century. Whether Earth can sustain this depends on consumption patterns, technology, and distribution.

The question isn’t just whether we can feed 10 billion but at what level and with what consequences. High consumption lifestyles have larger footprints. Climate change may reduce agricultural productivity. Ecosystems provide services beyond food. Carrying capacity is concept more than number — a reminder that growth encounters limits, even if those limits are complex and dynamic.

Go Deeper

Books

  • The Limits to Growth by Meadows, Meadows, Randers & Behrens (1972) — The famous Club of Rome report. Controversial, influential, surprisingly accurate in some predictions.
  • Collapse by Jared Diamond — Case studies of societies that exceeded carrying capacity: Easter Island, the Maya, Norse Greenland.
  • An Essay on the Principle of Population by Thomas Malthus (1798) — The original argument about population outstripping resources.

Essays

  • Garrett Hardin’s “The Tragedy of the Commons” (1968) in Science — How shared resources get overexploited when individual incentives diverge from collective welfare.
  • Verhulst’s 1838 paper “Notice sur la loi que la population suit dans son accroissement” — The original logistic growth model.

Related: ecology, ecological succession, fat tails